The manga is much more plot-driven, Tsukune is still everyone's love interest but I never felt that it was fan-service over content the way the anime was. From what I remember about the anime it was pretty much always: Moka transforms (in a fanservice-heavy sequence) and squashes the villain of weak in seconds. In the manga the villains gradually get stronger, team-work between the girls becomes more necessary and even Tsukune has to start finding ways to power up, rather than remaining the eternal damsel-in-distress. You start getting plots where the stakes are fate-of-the-world or at least fate of one member of the group being serious enough for the rest of the girls to set aside their rivalry to go to the rescue of whichever character is taking their turn at being in distress.
Ruby ends up being the designated fan-service character, but it's usually played for laughs: Ruby's personality is the most conservative of the girls, she's barely even a real member of the harem, more like an adviser/older sister to the group... Then she suddenly shows her hidden S&M leanings to everyone's shock.
The manga is very much superior to the anime IMO



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